Renée Veenstra successfully defends PhD on godwit foraging ecology

Sensing the soil

We are proud to celebrate another milestone in the journey of one of our young researchers. On 23 June 2026, Renée Veenstra successfully defended her PhD thesis at the University of Groningen under the Godwit Landscape Project. Started in 2021, the Godwit Landscape Project uses black-tailed godwits as indicators of the ecological health of Dutch dairy-farming landscapes. Within this framework, Renée investigated the foraging ecology of adult godwits on their breeding grounds.

Although godwits mainly feed on earthworms, researchers knew little about their prey preferences and foraging conditions. Using field observations, DNA metabarcoding, soil sampling, and experiments, Renée revealed a more complex foraging ecology.

Following food beneath the surface

Renée examined godwit diets during two key periods: a. Pre-nesting period (shortly after arrival on the breeding grounds) and b. Post-breeding period (after breeding, before migration). She found a clear seasonal shift in diet, from earthworms in wet spring soils to cranefly larvae (leatherjackets) later in the season and sometimes on spiders, when drier, harder soils make earthworms increasingly difficult to access.

Illustration by Marijke Klamer. Diet shift of the black-tailed godwit from pre-nesting to post-nesting period, from worms to the larvae of the cranefly.

One of the thesis’s key findings was that godwits are selective feeders, preferring red earthworms even when grey earthworms are more abundant. This challenged the assumption that all earthworms are equally valuable and showed that prey quality, accessibility, and abundance shape foraging behaviour. Red earthworms may be easier to detect, more accessible, or nutritionally superior.

The importance of moist grasslands

A central theme throughout the thesis was the importance of soil moisture. Experimental work using geophones showed that moving earthworms generates stronger and more detectable vibrations in moist soils, particularly red earthworms. These findings support the idea that godwits use specialized sensory receptors to locate prey beneath the soil surface. Moist soils improve both prey accessibility and detectability.

Illustration by Marijke Klamer. Worms can produce detectable vibration in the ground, which helps Black-tailed godwit to find worms.

Field studies showed that traditionally managed grasslands support more red earthworms than intensively managed fields. Higher water tables and favourable soil conditions provide more reliable food resources throughout the breeding season, highlighting the importance of maintaining moist grasslands to sustain godwit populations in an increasingly drought-prone landscape.

Illustration by Marijke Klamer. Black-tailed Godwit feeds on crane fly’s larvae (leatherjacket).

Growing as a researcher

Renée’s interest in meadow birds grew from earlier work on soil fauna, decomposition processes, and lapwing conservation. Her PhD combined these interests to explore how below-ground ecological processes influence godwits. Along the way, she developed skills in research design and fieldwork coordination, while learning to cope with uncertainty. She credits the supportive BirdEyes and Godwit team for helping her grow as a researcher, especially through discussions and constructive feedback. ‘Being altogether on every Wednesday, the afternoon discussion on ongoing research was very nice for me because it helped to shape the next steps, and I think it should continue,’ said Renée.

Renée’s research revealed how prey behaviour, soil conditions, and grassland management shape food availability for godwits. Her findings can inform future conservation efforts, and she hopes to continue exploring other ecological aspects of meadow bird conservation.

Centre for global ecological change at the University of Groningen

BirdEyes is a science and creative centre that views the world - almost literally - through the eyes of birds. More and more birds are flying around with tiny transmitters, loggers and other high technology on their backs and legs. This generates an unimaginable amount of information. By cleverly combining such data with other sources of information, and by using new ways to tell stories and share the insights with, BirdEyes strives to open up a new knowledge network. The centre at the Faculties of Science & Engineering and Campus Fryslân aims to be an innovative part of the University of Groningen and is linked to the Rudolph Agricola School for Sustainable Development. BirdEyes, with empirical and inspirational roots in the farthest corners of the world.

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